Here's what caused black stains on Leonardo da Vinci's Codex Atlanticus
Briefly

Researchers at the Politecnico di Milan in Italy examined mysterious black stains on a folio of Leonardo da Vinci's Codex Atlanticus and confirmed the presence of starch and vinyl glues in the affected areas.The glues were most likely applied during an earlier restoration effort some 50 years ago, according to a recent paper published in the journal Scientific Reports.
Read at Ars Technica
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